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ARCTIC OCEAN - A multi-year ice floe slides down the starboard side of the Coast Guard Cutter Healy Aug. 12, 2009, as the ship heads north into even thicker ice. "You can tell that this is a multi-year ice floe by the light blue melt ponds that have formed on top of the floe," said Pablo Clemente-Colón, chief scientist at the U.S. National Ice Center. Photo Credit: Patrick Kelley, U.S. Coast Guard. ARCTIC OCEAN - A multi-year ice floe slides down the starboard side of the Coast Guard Cutter Healy Aug. 12, 2009, as the ship heads north into even thicker ice. "You can tell that this is a multi-year ice floe by the light blue melt ponds that have formed on top of the floe," said Pablo Clemente-Colón, chief scientist at the U.S. National Ice Center. Photo Credit: Patrick Kelley, U.S. Coast Guard

The technique of overseeding where new grass is added to the existing turf for thicker and healthier coverage

The technique of overseeding where new grass is added to the existing turf for thicker and healthier coverage

Gymnosperm Stem: Cork and Cortex in Three Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Annual Rings in Three Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Longer and thicker

Longer and thicker

Thank you

Gymnosperm Stem: Pith in Two Year Pinus

cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Wavy, zig-zag lines - Thinner and thicker versions. Irregular li

Wavy, zig-zag lines - Thinner and thicker versions. Irregular li

Gymnosperm Stem: Medullary Rays in Three Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Syrah Grapes on the Vine

Wine-making grapes, like these Syrah (or Shiraz) grapes, require different growing conditions than grapes meant for eating. Cultivating wine grapes involves trying to concentrate sugars and encourage thicker skins, because these are what give wine its flavor. To do that, one of the most important factors to look for is the drainage of the soils. These grapes are grown on a greenstone schist formation in Westminster, Maryland. Photo credit: Alex Demas, USGS. Watch a video about "The Science of Good Taste — Geology, Food, and Wine" at on.doi.gov/FoodWine You can also learn more about "Oktoberfest! The Science of Beer and Wine Making" at on.doi.gov/Oktoberfest. Wine-making grapes, like these Syrah (or Shiraz) grapes, require different growing conditions than grapes meant for eating. Cultivating wine grapes involves trying to concentrate sugars and encourage thicker skins, because these are what give wine its flavor. To do that, one of the most important factors to look for is the drainage of the soils. These grapes are grown on a greenstone schist formation in Westminster, Maryland. Photo credit: Alex Demas, USGS. Watch a video about "The Science of Good Taste — Geology, Food, and Wine" at on.doi.gov/FoodWine You can also learn more about "Oktoberfest! The Science of Beer and Wine Making" at on.doi.gov/Oktoberfest

Logarithmic spirals with thinner and thicker lines

Logarithmic spirals with thinner and thicker lines

Roti Bom flatbread

Roti Bom flatbread

Gymnosperm Stem: Phloem in Three Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Wavy, zig-zag lines - Thinner and thicker versions. Irregular li

Wavy, zig-zag lines - Thinner and thicker versions. Irregular li

Gymnosperm Stem: Two Year Pinus

cross section: Pinus stem magnification: 40x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 40x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Paper roses.
Paper roses made arrangement of paper. White, thicker paper decorative. Retro object. Paper roses.
Paper roses made arrangement of paper. White, thicker paper decorative. Retro object.

Gymnosperm Stem: Pith in Three Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Wavy, zig-zag lines - Thinner and thicker versions. Irregular li

Wavy, zig-zag lines - Thinner and thicker versions. Irregular li

Gymnosperm Stem: Epidermis and Cortex in Two Year Pinus

cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous.

Female white walking shoes with thicker straps

Female white walking shoes with thicker straps

Gymnosperm Stem: Cambium in Two Year Pinus

cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

ARCTIC OCEAN - A multi-year ice floe slides down the starboard side of the Coast Guard Cutter Healy Aug. 11, 2009, as the ship heads north into even thicker ice. "You can tell that this is a multi-year ice floe by the light blue melt ponds that have formed on top of the floe, " said Pablo Clemente-Colón, chief scientist at the U. S. National Ice Center. Photo Credit: Patrick Kelley, U. S. Coast Guard

Hydrangea pruning year 2

Hydrangea pruning year 2

Green ivy grows thicker on the wall

Green ivy grows thicker on the wall

Happy Senior man cutting tree with loppers.

Happy Senior man cutting tree with loppers.

Gymnosperm Stem: Cambium in Three Year Pinus

cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Two Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

A single drop of blood into a clear class of water. Blood may be thicker than water but there is always more water.

Kelsey Trail No. 3

As Kelsey Trail No. 3 nears Winter Cabin, the scratchy chaparral vegetation becomes thicker. Long sleeves and pats are highly recommended for the route between Dorsey Spring and Winter Cabin. Photograph by Deborah Lee Soltesz. Credit USDA Forest Service, Coconino National Forest. For more information about this trail, see the Kelsey Trail No. 3 trail description on the Coconino National Forest website.

Bird; a surname

Also called birds. Any of various groups of vertebrates having warm blood eggs, breathing with the lungs, almost all body feathers, walking hind limbs, and wings in the forelimbs. Also called birds. Any of various groups of vertebrates having warm blood eggs, breathing with the lungs, almost all body feathers, walking hind limbs, and wings in the forelimbs.

Gymnosperm Stem: Three Year Pinus

cross section: Pinus stem magnification: 40x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 40x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

The golden headed lion faced marmoset (Leontopithecus chrysomelas)

The golden headed lion faced marmoset (Leontopithecus chrysomelas)

Wavy, zig-zag lines - Thinner and thicker versions. Irregular li

Wavy, zig-zag lines - Thinner and thicker versions. Irregular li

Thinner and thicker wavy, zig-zag horizontal line elements

Thinner and thicker wavy, zig-zag horizontal line elements

Gymnosperm Stem: Three Year Pinus

cross section: Pinus stem magnification: 40x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 40x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Cortical Resin Duct in Three Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

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